Friday, 28 September 2018

Eastern Forest Outskirts


Most of the land to the south and west of the hot plains is covered in dense jungle and rainforest. However, much of the eastern parts of these forests have recessed as a result of Cursor activity. Although the climate should allow the area to be more densely forested, the species of Cursor that inhabits the mossland just east of forest prevents many trees from growing there, since they’re consumed as small shrubs as the animals graze. The change from jungle to mossland is gradual, with an area of open woodland interspaced with mossland between the two. It is the wildlife that inhabits this area that will be discussed here.


Virgata purpura





Size: 1.5 metres tall discounting tail

Diet: leaves, grass, sometimes fruit and seeds

Habitat: savanna, woodland outskirts

Colouration: purple, with stripes of light and dark purple and black

Symmetry: 6 fold radial symmetry

Reproduction: sexual, all hermaphrodites. Lays eggs.

These animals are one of many species belonging to the fairly large family Folivoridae, a lineage closely related to Muscivoridae. Like muscivorids, they have a long proboscis, but they are distinguished from the related group by a longer and thinner tail with a tip adapted for grasping food. Although their tails are their primary means of gathering food, their proboscises haven’t atrophied; they are still useful for meeting the tail half way after food is gathered, so they needn’t reach all the way to their mouth. Their proboscis also allows them to drink ammonia without having to bend down.

Virgata purpura prefer to inhabit more open spaces than most other folivorids, and engage in a greater extent of herding behaviour, comparable to that of muscivorids. They have a slight build with fairly long legs and are quite agile runners, allowing them to effectively escape from predators even in lightly forested areas. Their exoskeletons and feathers are patterned in stripes, which not only serves to break up their form but visually confuses predators when they’re moving quickly in large groups. Since they’re small enough that the largest Acuti species of the area (who possess colour vision) are able to hunt them, they are purple in colour in order to blend in with the purple vegetation around them.

Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
Kingdom: Hydrogenia
Phylum: Plurafistulata
Superclass: Exoskeletida
Class: Lactilinguida
Subclass: Plumalia
Order: Herbivora
Family: Folivoridae
Tribe: Velocini
Genus: Virgata
Species: V. purpura


Procera collaris



Size: 4.5 meters tall up to tail, 8 meters with tail stretched up

Diet: leaves and fruit

Habitat: open woodland, savanna

Colouration: bright green and blue patterns, circle of yellow around the base of the tail

Symmetry: 8 fold radial symmetry

Reproduction: sexual, all hermaphrodites. Lays eggs.

Like Virgata, this species also belongs to the family Folivoridae. However, the two species rarely run into competition, since Procera eat far higher leaves than Virgata do. Their long tails and legs allow them to reach the high and relatively untouched foliage of some of the tallest trees of the forest outskirts, almost completely ignoring those in reach of Virgata. In order to support their weight at such heights they have a relatively light build, with a proportionately short body in comparison to their leg and tail length. Without their proboscis they would be unable to reach such heights, as squatting down to reach ammonia to drink is impractical for them.

The species is too large to be hunted by acuti, and since their only natural predators are the colour blind venators they are able to use the colour of their feathers for display. However, although colourful, the shade of their feathers is patterned in such a way so as to break up their form from the point of view of any colour blind animal. This way, they are able to stand out to other members of their own species, with colouration distinct from the foliage around them, while at the same time being effectively camouflaged from their predators.

They have a social structure based on a similar system of grooming as Minopuniceus, but with the sharing of fruit among group members playing an additional role. They don’t always eat the food they gather straight away; while they will often eat leaves as they find them, when particularly large fruit is found they instead keep them in case they wish to trade it to win the favour of other members of the group. Because of this dominant members of a group will usually eat far more fruit. There is usually a single individual at the very top of the hierarchy, with the yellow feathers at the base of their tail particularly prominent and brightly coloured. This individual is most often also the tallest.

They are very intelligent animals, with more complex relationships than the otherwise socially similar Minopuniceus. As such they have much smaller group sizes than both Minopuniceus and the related Virgata, allowing them to keep track of the group dynamics and know everybody more intimately. They tend not to gather closely together, and do not possess the same herding instinct common among muscivorids and many folivorids. They simply don’t need to live in large groups, as their size offers enough protection on its own.

Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
Kingdom: Hydrogenia
Phylum: Plurafistulata
Superclass: Exoskeletida
Class: Lactilinguida
Order: Herbivora
Family: Folivoridae
Tribe: Procerini
Genus: Procera
Species: P. collaris



Eastern Tree Lethargid
(Ramiflagellum solum)




Size: 30 - 40 centimetres long on average

Diet: meat, often preys on animals larger than itself

Habitat: arboreal

Colouration: brown, reddish brown or purplish brown

Symmetry: bilateral, originally evolved from radially symmetrical animals

Reproduction: sexual, all hermaphrodites. Lays eggs.

Venators are a very large and successful class of animals, and although most are predatory they fill a large range of niches and inhabit a large range of environments. Not all venators focus on pack hunting and chasing down prey like Leolupus. Some, namely those belonging to  the family Lethargidae, are much smaller and instead have a much more solitary and cunning approach to hunting, which involves hiding and waiting for their prey to approach. They have very long lashing tongues with sharp barbs, and many species utilise toxins or bacteria to make up for their lack of size or strength.

Ramiflagellum solum is a lethargid much more adapted to climbing than other members of the family, spending the vast majority of its time in trees. Their genus is distinct from other members of the family by their smaller size and the presence of opposable, grasping digits on all six limbs, as well as a longer and more elastic tongue than most other species. While a small size is beneficial to terrestrial arboreal animals, it is even more so on Amthalassa. Many trees are lightweight and only supported by the lift they’re afforded by sacks of hydrogen, so if any animal is large enough to overcome this lift on a given branch the branch will collapse. Because of this, members of the genus Ramiflagellum are comparatively weak and lightweight even compared to other related groups, so they have to rely even more on their ability to hide.
When a prey animal is sighted, Ramiflagellum solum will lash its tongue out from the tree concealing it, using its well-developed senses and accuracy to target weak spots or gaps between exoskeleton sections. If the strike is successful, and the teeth pierce the animal’s flesh, a form of modified, toxic stomach acid that is kept on the tips of its teeth is injected into the animal’s blood. This venom will slowly weaken the animal, and Ramiflagellum will stealthily follow it until it eventually falls unconscious and becomes easy to kill. This way, Ramiflagellum is able to take down prey far larger than itself.
Although primarily arboreal, they spend more time on the ground than their cousins deeper into the rainforest further west, and lack many of their adaptations such as a prehensile tail.

Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
Kingdom: Hydrogenia
Phylum: Plurafistulata
Superclass: Exoskeletida
Class: Venatores
Order: Insectilupiformes
Family: Lethargidae
Genus: Ramiflagellum
Species: R. solum


Woodland Septacaud


(Ramicauda orientalis)





Size: 60 centimetres to 1 metre

Diet: fruit and leaves

Habitat: arboreal

Colouration: purple

Symmetry: radial (seven fold)

Reproduction: sexual, all hermaphrodites. Lays eggs lacking hard shells in bodies of ammonia.

These animals inhabit the wetter and more densely forested part of the area, spending most of their adult lives high in trees but returning to the ground to lay eggs. These eggs are laid in ponds, where the aquatic, snake-like larvae will spend all their time until they’re developed enough to survive on land.

Above: Ramicauda disguising itself on a tree branch
Like most Septicauds, Ramicauda can use hydraulic pressure to stretch their tentacles to several times their resting size, making them well adapted to tree swinging in their arboreal habitat. They also have the ability to change their shape slightly by adjusting the hydraulic pressure under their skin, allowing them to effectively disguise themselves from predators. This ability is also used for signalling within their species.
Above: A young Ramicauda in its aquatic larval state



Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
 Kingdom: Hydrogenia
 Phylum: Plurafistulata
 Superclass: Multicaudata
 Class: Septicaudata
 Order: Polymorpha
 Family: Ramicaudidae
 Genus: Ramicauda
 Species: R. orientalis






 Hinnuleus occidentalis


Size: 90 centimetres on average, discounting tail

Diet: leaves, moss, fungus, sometimes fruit and seeds

Habitat: woodland

Colouration: red, purple and brown, usually in a pattern that breaks up their form

Symmetry:  8 fold radial symmetry

Reproduction: sexual, all hermaphrodites. Lays eggs.

Hinnuleus is a group of folivorids much smaller than Virgata purpura, preferring to make use of their small size to hide in the dense foliage of the woodlands at the expense of running ability. They are very well camouflaged, but their small size doesn’t allow them to reach high leaves. Instead they will usually focus on leaves closer to the ground, although this is more of an advantage than anything else; with Hinnuleus, Virgata and Proceri focusing on leaves at three different heights, they are all able to occupy the same woodland in spite of their similar diets without running into competition.

Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
Kingdom: Hydrogenia
Phylum: Plurafistulata
Superclass: Exoskeletida
Class: Lactilinguida
Subclass: Plumalia
Order: Herbivora
Family: Folivoridae
Genus: Hinnuleus
Species: H. occidentalis


Duocaputina
(Cornucerva magna)



Size: 1.4 to 1.7 metres long

Diet: primarily fungi, sometimes shrubs, moss or kelp tree leaves

Habitat: woodland

Colouration: reddish brown

Symmetry:  biradial

Reproduction: sexual, all hermaphrodites. Lays eggs.

There are multiple occasions on Amthalassa where initially radially symmetrical animals have become at least partially bilaterally symmetrical. This usually happens in faster animals, so they can become more specialised at running in a single direction. Duocaputina is one such case. This group is actually a member of the family Folivoridae, but have changed much from the basal body plan of the group. To say they’re bilaterally symmetrical isn’t entirely accurate; they’re biradially symmetrical, with two distinct lines of symmetry rather than one. They have a distinct top and bottom, and sides of the body distinct from the front and back. However, the front and back of the body are identical, with one “head” on either end. This is probably advantageous in that the animal can begin running in either direction upon sighting a predator, without having to turn before running as much as they’d otherwise have to if they were purely bilaterally symmetrical.

The animal’s mouth is underneath its body. Since their tail isn’t as long as that of most other folivorids, as it has been reduced in size to make the animal more aerodynamic for running, they rely on their proboscis for feeding. Because of this, they are only usually able to eat food off the ground, so their diet primarily consists of fungi growing on the forest floor. However, they will occasionally use their weight to pull down smaller kelp trees by walking over them, allowing them to reach their leaves.

The genus Cornucerva is distinguished from other members of the tribe Duocaputina by the presence of horns, as well as a particularly thick, shell-like exoskeleton on the back and “heads”. They use these horns to settle in-group disputes, as their primary means of intraspecies fighting is head-butting. Sexual selection also played a role in the development of horns.

Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
Kingdom: Hydrogenia
Phylum: Plurafistulata
Superclass: Exoskeletida
Class: Lactilinguida
Subclass: Plumalia
Order: Herbivora
Family: Folivoridae
Tribe: Velocini
Subtribe: Duocaputina
Genus: Cornucerva
Species: C. magna


Caudarostrum externum





Size: 20 - 50 centimetres, not including tail

Diet: seeds, leaves and fruit

Habitat: arboreal

Colouration: purple

Symmetry:  8 fold radial symmetry

Reproduction: sexual, all hermaphrodites. Lays eggs.

Although most folivorids use their tails graspers to gather food, they are usually very poor at biting up food with it. However, there is a branch of Folivoridae that have developed hard beaks at the end of their tails. Although the ancestral form was likely a reasonably large herbivore, most extant species are much smaller than is the norm for folivorids, and many, such as Caudarostrum, are even tree dwellers.

Caudarostum spends almost all of its life hanging from trees with its hook-like claws as it browses for food with its beak. Its beak is particularly well adapted for eating seeds, which make up a large portion of its diet.

Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
Kingdom: Hydrogenia
Phylum: Plurafistulata
Superclass: Exoskeletida
Class: Lactilinguida
Subclass: Plumalia
Order: Herbivora
Family: Folivoridae
Subfamily: Caudarostrinae
Tribe: Caudarostrini
Genus: Caudarostrum
Species: C. externum

Fresh Ammonia Fish



Many species of Amthalassan fish inhabit the ammonia rivers and lakes of the woodland. Most plurafistulate fish on Amthalassa are long and snake-like in appearance, or somewhat resemble an eel. Unlike Earth fish, most lack movable fins, and many lack fins altogether, although a number of fish species possess facial tentacles. Those with tentacles belong to the clade Ouropsia (although many branches of Ouropsia have lost their tentacles), and it is this group that most land plurafistulates descended from (and in fact, still belong to). 



Magnicollum pondus



Size: 2 metres tall, discounting tail

Diet: leaves, sometimes fruit

Habitat: open woodland

Colouration: bright green and red

Symmetry: 8 fold radial symmetry

Reproduction: sexual, all hermaphrodites. Lays eggs.

Most folivorids make use of their long tails to reach high vegetation. Although Magnicollum, too, focuses on higher foliage, it does not use the same method as its relatives. While on Earth, trees are fairly rigid and difficult to manipulate, this is not the case with kelp trees. Magnicollum is much stockier for its height than other folivorids, and makes use of its high weight to anchor itself to the ground as it pulls down kelp trees with its muscular “neck”. As soon as the animal obtains the vegetation it needs, it releases its grip on the tree and allows it to float back up to its original height.

There is a limit on how large a tree an individual can pull down this way; larger trees have more hydrogen sacks, and so are more buoyant, and will remain floating under the weight of a larger animal. This is why members of the genus are so large, otherwise they cannot effectively pull larger trees down.

Much of the animal’s extra weight consists of exoskeleton. Since this is fairly unenergetic tissue, this means the animal can be much more massive without requiring more energy, so the full benefits of being able to obtain more food by pulling down larger kelp trees can be felt. This thick exoskeleton, coupled with the animal’s large size, makes them well protected from predators. Because of this, as well as the fact their large size makes camouflage difficult, their colouration is mainly for display purposes.

Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
Kingdom: Hydrogenia
Phylum: Plurafistulata
Superclass: Exoskeletida
Class: Lactilinguida
Subclass: Plumalia
Order: Herbivora
Family: Folivoridae
Genus: Magnicollum
Species: M. pondus

Claviungula corticis



Size: 20 centimetres from toes to tail tip

Diet: meat, tube bugs

Habitat: arboreal

Colouration: brown, reddish brown or orange-brown exoskeleton. Feathers also tend to be dull colours to camouflage them from colour seeing prey.

Symmetry: 8 fold radial symmetry

Reproduction: sexual, all hermaphrodites. Lays eggs.

These are acuti belonging to the family Sciuracutidae, a group of very small, tree dwelling acuti. Most sciuracutids, Claviungula included, climb by digging their sharp, long claws into the trunks of wood trees, a means of locomotion where lower body weight provides an obvious advantage.

Like most acuti, sciuracutids tend to be carnivorous. Due to their size, however, they tend to go after very small prey, much smaller than even other acuti. Claviungula has a diet composed primarily of tube bugs and grassbugs, but will also eat small plurafistulates, including young Caudaerostrum on occasion. Their primary means of hunting plurafistulates is by sneaking up on them and striking at the last minute, but they usually simply lash their clawed tongues out to eat tube bugs or use their long claws to pick them out of trees.

Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
Kingdom: Hydrogenia
Phylum: Plurafistulata
Superclass: Exoskeletida
Class: Lactilinguida
Subclass: Plumalia
Order: Acuta
Family: Sciuracutidae
Genus: Claviungula
Species: C. corticis


Tree Grazing Floater
(Foliovis aculeatus)
Size: up to 10 metres long


Diet: leaves

Habitat: forests, above trees

Colouration: brown, orange-brown, red-brown

Symmetry:  bilateral

Reproduction: sexual, all hermaphrodites. Lays eggs.

Like other floaters, those in the family Arboripascidae have a symbiotic relationship with hydrogen producing plants, allowing a large hydrogen sack to grow on their bellies to remain buoyant. Arboripascids are far larger than the faster and more flexible rectabullids, plodding along very slowly.

Foliovis spends its time above the highest canopy of forests, grazing the top layer of leaves as another animal might graze grass. They spend much of the day eating, and possess three proboscises that allow them to forage for food at a fast rate.

If they depended just on their mouths, they’d have to spend a lot of energy moving about just to obtain the large quantities of food needed to support their size, and for a creature this large and slow this isn’t very practical. They’d use more energy than they’d obtain. However, they have three fast moving and long proboscises – each formed by the fusion of two tongues – where they consume almost all of their food from.

They spend a lot of time hovering in a single place, eating as much of the leaves as they can while moving only their tongues, before moving on to another area.

In order to support their large size, their balloon leaf cavity takes up most of their body. To further decrease their weight, they have many air sacks throughout their body that only allow hydrogen from the balloon leaf in.

Arboripascids are warm blooded and capable of localised heating in different parts of their body. To further lower their density, their hydrogen sacks are heated a great deal more than the rest of their bodies. The rest of the body is kept cooler, at temperatures more optimal for their physiology. This heating lowers the density of the hydrogen, contributing to the buoyancy of these animals.

Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
Kingdom: Hydrogenia
Phylum: Plurafistulata
Superclass: Exoskeletida
Class: Folivesicata
Subclass: Vesicaspecata
Order: Aerobalaenacea
Family: Arboripascidae
Genus: Foliovis
Species: F. aculeatus


Woodsnake
(Silvavora unguis)

Size: 40 to 80 centimetres long

Diet: wood, plant exoskeleton, leaves

Habitat: trees

Colouration: dull brown or reddish brown

Symmetry: 8 fold radial

Reproduction: sexual or asexual, all hermaphrodites and can self-fertilise. Lays eggs.

A distant relative of the earthsnake, woodsnakes share their relative’s powerful beaks, but use them to bite into wood or plant exoskeleton rather than soil. Since they aren’t burrowing animals, they lack the lung capacity of earthsnakes, and the facial appendages used for digging by their relatives can do nothing more than manipulate food.

Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
Kingdom: Hydrogenia
Phylum: Plurafistulata
Superclass: Exoskeletida
Class: Sectatestae
Order: Bidigiti
Family: Silvavoridae
Genus: Silvavora
Species: S. unguis

Haploformica dumus




Size: gametophytes are tiny, measuring from under a millimetre to a centimetre depending on the type. Sporophytes grow throughout their whole lives and can range from 15 cm to several meters, though are usually quite small, about a meter in height.

Diet: plants and small tube bugs brought back by gametophytes

Habitat: woodland, savanna

Colouration: gametophytes are often dark and dull in colour for camouflage. Sporophytes have grey bark and orange leaves.

Symmetry: gametophytes have radial symmetry. Sporophytes have roughly radial symmetry, with branching.

Reproduction: reproduces sexually with gametophytes

This is one of the many species of Amthalassa belonging to the mostly mobile group of plants. Many mobile plants, however – especially those belonging to the class Immota – are sessile during certain stages of their life, planting their roots into the ground permanently once they find a suitable location.

Like many other mobile plants, the gametophytes of Haploformica are capable of locomotion. The order it belongs to, Rapidasperma, is unique in that the role of these gametophytes aren’t limited to reproduction; they also defend the plant, do work for the plant, and gather food for it from its surroundings. They produce gametophytes almost all the time, and in fact most gametophytes are unable to perform their reproductive function.

Haploformica produces various different specialised gametophytes for different functions. The majority of gametophytes are workers, spending most of their time gathering food. They also produce gametophytes specialised for defence, able to attack any animal threatening the plant, and they can send out gametophytes that act as scouts.
A sample of specialised gametophytes

Haploformica trees are also able to produce male gametophytes, which are able to travel great distances to other Haploformica plants to merge with any female gametophytes inside. Any female gametophytes these plants produce never leave them, and unlike other gametophytes they can’t move on their own. Male gametophytes have a feathery appendage on their backs that allows them to be blown in the wind to cover greater distance, which they can also pull down if they wish to drop or stay on the ground.

When a tree grows a seed, this seed is then taken by a worker gametophyte to a suitable location. Unlike many other species in the class Immota they are much less active in their early years, once they start growing from their seed. Other immotans are active at first before finding a suitable location, then remain sessile.

The trees control their gametophytes with pheromones, and gametophytes also interact with each other this way to an extent. Like most mobile plants they do possess very rudimentary intelligence, in spite of seeming like an ordinary sessile plant, so they’re able to recognise danger and command their gametophytes in response to their surroundings. They even possess simple eyes. Their gametophytes are also capable of limited cognition, but to a far lesser extent than the trees themselves.

Taxonomic classification
Tree: Amthalassavitae
Domain: Parvigrana
Kingdom: Ramipedes
Phylum: Magnisemina
Class: Immota
Order: Rapidasperma
Family: Haploformicaceae
Genus: Haploformica
Species: H. dumus


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